The game opens for both book one and two with Toad and Toadette getting the Power Star at the top of the building, until the villainous giant crow named Wingo takes the star and kidnaps either Toad or Toadette. As the game progresses, Toad and Toadette defeat Draggadon, the king of Pyropuff Peak. At the end of book two, Wingo kidnaps Toadette and sends Toad flying away from the battle tower.
In book three, Wingo leaves the cave with the star, before Toadette escapes and attempts to reunite with Toad. As Wingo kidnaps Toadette again and goes to the watch tower, Toad climbs up to it filled with treasures and throws the summoned giant turnip at Wingo who swallows it for heavy weight and falls from the sky, freeing Toadette and the star. At the end of book three, Toad rushes off alone, pursues the falling Green Star in the glass pipe and enters it, setting the stage for Super Mario 3D World.
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